Chapter 19 Evolution
Philosophers and scientists have been busy to solve the riddle as to how the universe and our earth were formed and how and when 'life' originated on earth. The branch of life science for the study of 'Origin of life' and evolution of different forms of life on earth was called Bioevolution or Evolutionary Biology by Mayer, (1970).
The study of universe or cosmos is called Cosmology. Our earth belongs to the Solar system having nine stars called planets constantly rotating around a common Sun. On the basis of the order of the distance from the sun these planets include Mercury, Venus, Earth, Mars, Jupiter, Saturn, Uranus, Neptune and Pluto while moon is a satellite of earth. The universe is made up of matter and energy and it was formed about 10 to 13 billion years ago as a red hot, dense, rotating gaseous cloud of cosmic dust called Ylem or primaeval matter. The Ylem consisted of particles of matter (like neutron, proton and electron) and antiparticles of antimatter. Scientists like Lemaitre (1931), Gamow (1948), Dicke (1964) etc. supported the Big-Bang Hypothesis which explains that collision between these particles and antiparticles caused a tremendous explosion to form atoms of hydrogen. Cosmic evolution began with the fusion of hydrogen atoms with progressively heavy atoms of different elements. Stellar systems and stars were formed by spreading of original gaseous cloud into the space and divided into smaller and larger masses. Most of the stars are masses of red hot gases even today.
Kant (1755) and Laplace (1796) supported Nebular Hypothesis which explains our solar system to have evolved about 4.5 to 5 billion years ago from a rotating red-hot gaseous cloud containing millions of free atoms of different varieties. First of all sun was formed when this cloud condensed, next the planets were thrown off from the sun and later on, in turn satellites were thrown off by planets. However, Nebular hypothesis was rejected by astronomers of present century like Weizsacker (1944), Alfen (1950) and Hoyle (1955).
According to them the sun was originally surrounded by a disk of rotating gas. Later many rotating concentric whorls were formed by break up of the disc and by gravitation and rotation particles of each whorl collected and condensed to form a planet. Our earth was formed about 4.6 billion years ago as a red hot gaseous cloud of free atoms with temperature of 5000o to 6000oC.
Structure of present earth. The earth is orange like in shape. Its pole to pole diameter is 12640 km and equatorial diameter is 12783 km. It is about 15
crore km away from sun and about 484000 km away from moon. Due to tremendous temperature gases existed in atomic form but gradually they cooled down in hundreds and millions of years into molten core. According to density other elements got stratified. Earth contains the central solid core, the middle mantle and shell and outer crust. The earth rotates at its polar axis in one day and it rotates
around the sun in 365 1
2
days or one year.
Life is the part and parcel of the universe and both are very intimately associated with each other. We know that “Life is the most unique, complex organisation of molecules, expressing itself through chemical reactions which lead to growth, development, responsiveness, adaptation and reproduction” that matter has achieved in our universe. Origin of life is a unique event in the history of universe.
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Francesco Redi (1668) showed that maggots could not be created from meat. Actually, the smell of meat attracts flies which lay eggs on the flesh. These eggs hatched into flies.
Spallanzani (1767) showed that even primitive, unicellular organisms cannot arise from non-living matter.
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Louis Pasteur also, used swan-necked flasks whose long, curved necks permitted exchange of air between outside and inside of the flask, but dust and bacteria were trapped along the wall of the neck. On tilting the flask, the bacteria got washed down into the broth, so that the latter became cloudy due to bacterial growth.
planets small units called ‘spores’. The spores were covered by a thick protective covering. When the spores got favourable conditions and temperature, the spore coat was dissolved and gave birth to initial living organisms. This theory does not explain as to how the life originated in space and how the life originated in spores remain impenetrable by ultraviolet and gama rays.
According to this theory, the Earth originated about 4,500 million years ago. When the earth was cooling down, it had a reduced atmosphere. In this primitive atmosphere nitrogen, hydrogen, ammonia, methane, carbon mono-oxide and water were present. Energy was available in the form of electric discharges by lightening and ultraviolet rays. As soon as the earth crust was formed, it was very much folded. Torrential rains poured over the earth for centuries and were deposited in deep places.
The atmospheric compounds, inorganic salts and minerals also came in deep places oceans, these molecules gave rise to a variety of compounds and finally to the self-duplicating molecules. Ultimately these molecules were enclosed in membranes derived from lipids and proteins, along with water and chemical compounds, giving rise to cell like units. Again random combinations may have led to the formation of chlorophyll– containing organisms which could produce their own food (autotrophs) by a process called photosynthesis. These organisms had a better chance to live because
they synthesise starch from carbon dioxide and water in presence of
sunlight. Starch could be used as further source of energy. During photosynthesis, oxygen was produced. The oxygen was used by
According to Oparin’s hypothesis–Origin of cells from coacervates
other organisms for respiration. Also oxygen, when acted upon by ultraviolet rays, formed ozone layer through which ultraviolet rays cannot pass. This layer is formed about 25 km. from earth's surface. After the formation of ozone layer, organisms could come to the surface of the ocean and could survive even on land, if thrown out of oceans. The Oparins's and Haldane's theory of origin of life is most accepted these days as it is supported by Miller's experiment duly supported by David Buhal, Melvin Kelvin's experiment etc.
Miller's Experiment : An American scientist (Biologist) Stanley Miller (1953) performed an experiment under support Oparin's theory of origin of life. He believed that basic compounds which are essential for life can be
synthesised in the laboratory by creation in the laboratory, on a small scale, the conditions which must have existed at the time of origin of life on earth.
Miller took a flask and filled it with methane, ammonia and hydrogen in proportion of 2:1:2 respectively at 0°C. This proportion of gases probably existed in the environment at time of origin of life. This flask was connected with a smaller flask, that was filled with water, with the help of glass tubes. In the bigger flask, two electrodes of tungsten were fitted. Then a current of 60,000 volts was passes, through gases containing bigger flask for seven days. At the end of seven days, when the vapours condensed, a red substance was found in the U-tube. When this red substance was analyzed, it was found to contain amino acids, Glycine and nitrogenous bases which are found in the nucleus of a cell.
From the above theory we conclude that life first originated in water. Therefore, water still continues to be an essential constituent of life.
The entire process of the origin of life, as proposed by Oparin, can be summarised as under –
The scientists have found that simple unicellular organisms (resembling modern cyanobacteria) were present on this earth about 3600 million years ago. It is believed, therefore, that life must have originated on this earth about 4600 to 3600 million years ago.
It must be clear that the earth's atmosphere at that time was quite different from as it exists today. The earth's atmosphere at that time was reducing, not oxidizing (as it is today). The primitive atmosphere of earth had hydrogen, nitrogen, water vapours, carbon dioxide, methane and ammonia abundance. Oxygen was not available in free state in sufficient quantities.
These compounds were formed at the time when sunlight could not reach earth because of dense clouds in the sky. Under such circumstances, the energy required for the synthesis of above–mentioned chemical substances must have been obtained from the cosmic rays and lightening in the sky. Even today, NH3 and other compounds are formed in the atmosphere during electric discharges. Thus, it is clear that all the amino acids required by living organisms could be formed in the condition of primitive atmosphere. However, this type of synthesis requires reducing atmosphere because the substances produced through these processes get destroyed by oxidation on coming in contact with oxygen. Since free oxygen is available in the atmosphere today, present day atmosphere is no longer reducing–it has turned oxidizing. Therefore, in the present circumstances, such a synthesis of chemical substances is not possible.
In fact, the present day organisms synthesise their complex molecules from simple organic substances with the help of enzymes only. However, enzymes themselves are proteins. Therefore, first of all protein must have been synthesized without the help of any enzyme.
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Other Metals
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and electric discharge
Oparin believed that the formation of protein was a very important step in towards the origin of life. Due to their Zwitterionic nature, the protein molecules. This enabled these colloidal structures to maintain their identify inspite of being surrounded by water molecules–forming a type of emulsion. The coalescence of these colloidal structures led to the formation of structures called coacervates. These coacervates had the ability to exchange substances with the surrounding water and accumulating required substances within them.
Sydney F. Fox's experiment : Sydney F. Fox of Florida University, obtained some complex molecules by heating upto 90°C a dry mixture of many amino acids found in living organisms. The molecules so obtained very much resembled the proteins. He heated these molecules in water and allowed the mixture to cool down. In the fluid so obtained, he could observe minute structures resembling the cells. He called them microspheres. The microspheres are surrounded by membranes, and these also reproduce vegetatively just like years. Surprisingly the biochemical processes like breakdown of glucose also occur their. However, electron microscopic examination of these does not reveal any cellular structure. On keeping in distilled water, these become turgid, but these get shrunk if immersed in salt solution. Oparin's coacervates and Fox's microspheres are infact, similar structures.
Earliest organisms : Evidences available so far indicate that the cells of the earliest organisms did not contain either nucleus or cell organelles. The molecules of nucleic acid were surrounded by a colloidal mixture (may be called protoplasm) of proteins and organic compounds. This, in turn, was surrounded by a thin membrane.
Water and soluble substances would pass through this membrane. Such cells which lacked nuclei were called prokaryotic cells. In 1966, some fossils have been discovered from 300 million years old rocks. These fossils are of prokaryotic organisms. Earliest organism is chemoheterotrops.
At such a time, some of these organisms developed the capability of synthesizing organic substances. Such organisms began synthesizing energy– giving substances (carbohydrates) from simple inorganic substances abundantly available in the environment. In this way, evolution of autotrophs from heterotrophs took place.
This was the beginning of autotrophic nutrition. However, it was quite different from the photosynthesis which is carried out by green plants, because it utilized energy obtained by anaerobic respiration (not solar energy). Therefore, such type of nutrition is also called chemoautotrophic nutrition. Such type of nutrition is observed even
today in the sulphur bacteria.
6 CO2 + 12 H2S ¾¾Ferm¾ent¾at¾ion® C6 H12O 11 + 6 H2 O + 12 S
Chemical energy
At the same time, from different chemicals present in the oceanic water evolved porphyrins which where like modern chlorophyll led to the evolution of present chlorophyll, so that these cells started utilizing H2 O instead of
H2S for photosynthesis.
6 CO2 + 12 H2S ¾¾Sol¾ar ® C6 H12O6 + 6 H2 O + 12 S
Energy
Till then, oxygen was not freely available in the atmosphere. However, gradually molecular changes in the bacteriocholorophyll led to the evolution of present chlorophyll, so that these cells started utilizing H2O instead of
H2S for photosynthesis.
6 CO2 + 12 H 2 O ¾¾Sol¾ar ® C6 H12 O6 + 6 H 2 O + 6O2
Energy
In this way, the prokaryotic cells which were chemoautotrophs, became photo autotrophic. These cells resembled modern cyanobacteria. In 1968, the forms of such types of cells have been recovered from 320 million years old rocks. These have been given the name Archaeospheroides barbertonensis. Due to the absence of well- defined nuclei in them, these have been included under the kingdom ‘Monera’. Thus, release of O2 in the atmosphere and its free avilability was the result of photosynthesis. This was a revolutionary change which greatly affected the course of organic evolution.
types of tissue combined to form special organs. From the organs, organ systems and ultimately complex bodies of organisms were formed.
The following are the evidences in favour of Organic Evolution :
(i) Evidences from Classification (ii) Evidences from Comparative Anatomy
(iii) Evidences from Physiology (iv) Evidences from Serology
(v) Evidences from Embryology (vi) Evidences from Palaeontology
(vii) Evidences from geographic distribution (viii) Evidences from Genetics
The homologous structures give us the answer that how the two different types of animals or organs have the same origin and functions to perform; this shows that they have changed themselves according to their different needs.
functional in allied animals. Such useless organs are termed as vestigeal and are of frequent occurrence. In human body alone, there are as many as ninety such organs. For example, vermiform appendix is vestigeal in man but functional in rodents, horse and other herbivorous animals.
Take the example of thyroxin hormones, which are similar in all
metabolism. The thyroid of human beings can be very easily replaced by the thyroid of cattle without any ill-effects and so on.
Different vestigial organs in man
Haeckel, a German biologist, gave a theory “Ontogeny repeats Phylogeny”. An individual organism during its development (ontogeny) tries to repeat the history of its race by different stages (phylogeny). The vertebrate development confirms the above statement. For example, the embryos of all vertebrates pass through a gilled stage. In fishes, the gills are present in the adult condition. In amphibians, gills are present in some forms of frog and in
tadpole. While in reptiles, birds and mammals these gills are never functional but nevertheless they are always present in the embryo.
Haeckel, believed that all those stages, which have occurred during the course of evolution of a particular animal, also pass through its development.
There are objections to this theory, but is throws enough light to show the interrelationship of animals.
A number of fossils have been discovered from time to time. Some of these fossils are very prefect in their state of preservation. For example, the fossils Archaeopteryx shows the characters of reptile on the one hand, and the characters of birds on the other hand, meaning that the birds have evolved from reptiles, the Archaeopteryx being a connecting link.
In the same way all the evolutionary stages of horse, elephant, camel and man etc., can be constructed. The earliest horse was known as Eohippus. It was eleven inches in height and made its appearance in Eocene time from some unknown five-toed ancestor. The Eohippus, after various evolutionary stages, transformed itself into the present-day horse Equus, which shown many dissimilarities from its great grand, grand ancestor. The various stages in its evolutionary life had been preserved as fossils. The same is the case with camel, elephant and man, etc. So this science of paleontology helps in a great deal in understanding the process of evolution.
Era |
Age million years |
Period |
Epoch |
Dominated Fauna |
Cenozoic (Age of Modern life) |
0.1 |
Quaternary |
Recent (Holocene) |
Modern man, mammals, birds, fishes, insects. |
2.5 |
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Pleistocene |
Extinction of grate mammals. Primitive man common. Evolution of human society & culture. |
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7 |
Tertiary |
Pliocene |
Evolution of primitive man–like forms from man like apes. Formation of modern mammals. |
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2.5 |
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Miocene |
Mammals at peak. Evolution of man–like apes. |
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3.8 |
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Oligocene |
Decline of egg laying and marsupial mammals. Rise of first monkeys and apes. |
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5.4 |
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Eocene |
Diversification of placental mammals. |
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6.5 |
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Paleocene |
Rise of first primates, placental mammals and modern birds. |
Mes ozoi c (Era |
135 |
Cretaceous |
Extinction of dinosaurs and toothed birds. Rise of first modern birds. |
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195 |
Jurassic (Age of Giant Reptiles) |
Origin of advanced lizards, crocodiles, alligators, marsupials toothed birds. Dinosaurs became large. Reptiles dominant. |
240 |
Triassic |
Origin of dinosaurs & primitive mammals (egg–laying mammals). Extinction of primitive amphibians. |
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Paleozoic (Era of Ancient life) |
285 |
Permian |
Extinction of many marine invertebrates like trilobites. Rise of modern insects. Evolution of mammals like reptiles. |
375 |
Carboniferous (Age of Amphibians) |
Origin of reptiles and winged insects. |
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420 |
Devonian (Age of fishes) |
Origin of first land vertebrates (amphibians) Fishes abundant. |
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450 |
Silurian |
Origin of jawed fishes and wingless insects. |
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520 |
Ordovician (Age of invertebrates) |
Origin of chordates with first jaw– less fishes (origin of vertebrates). Invertebrates abundant. |
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570 |
Cambrian |
All invertebrate phyla established. Trilobites (swimming crustaceans which do not exist today) dominant |
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Proterozoic (Era of early life) |
2300
3600
3800
4600 |
Pre–cambrian |
Origin of marine metazoans including sponges, cnidarians, annelids, molluscs and arthropods. Scanty fossils. Origin of prokaryotes (Monera) and Eukaryotes (Protista). Origin of life. |
Archeaozoic (Era of invisible life) |
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No life? |
1. Peripatus (Arthropoda) |
2. Limulus (Arthropoda) |
2. Nautilus (Mollusca) |
4. Neopilina (Mollusca) |
5. Lingula (Brachiopoda) |
6. Latimeria (Coelacanth fish) |
7. Sphenodon (Reptilia) |